I can't believe anyone would think to fill a water-tank BEFORE they set off! Not only does it make the caravan colossally heavy, but it is extremely dangerous, making the outfit highly-unstable and likely invalidating your insurance! You should always drain the tank completely before towing.
The main advantage of an onboard tank is that you have more water stored right at the van (very helpful on a site without a shower/toilet block, for example!), and to keep the temperature of the water above freezing if you tour in the winter. I don't know about keeping it cooler during warm weather but I can see that it might do this, too.
It's very easy to fill/use; just 'plug' the connector hose from the aquaroll in exactly the same way you normally would, and then open an additional tap inside the caravan (in my case, under the front chest) to fill the tank. One aquaroll = approx one full tank. You can then refill the aquaroll and have it ready to top up the tank, if you want. It isn't necessary to fill the tank fully to use it (the caravan's control display will tell you how much water is in the tank), or even use the tank at all - you can set the internal tap to feed water directly from the aquaroll to the caravan, bypassing the tank altogether (if you're only having an overnight stop, for example).
Regards, Abi